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Dr. Richard M. Engeman

Dr. Richard Engeman is a biologist and research biometrician at the NWRC working on the monitoring, indexing, and the economics of control for a variety of species. His areas of expertise include statistics, experimental design, population monitoring, distance sampling, and plotless density estimation.

NWRC Research Project: Avian and Invasive Species Population Management

Current Research

  • Feral swine population monitoring, damage estimation, valuation, and disease spread
  • Florida invasive reptiles, especially pythons, iguanas, monitor lizards
  • Gambian giant pouched rat management
  • Population indexing method development
  • Damage estimation research
  • Bioeconomic valuations and analyses
  • Population monitoring for international collaborations
  • Economics of control to protect endangered species


  • Ph.D., University of Colorado Health Sciences Center, Biometrics, Sinusoidal input design for model discrimination and parameter estimation
  • M.S., Colorado State University, Statistics, Generalized least squares and two-step estimators of the parameters of the Weibull distribution
  • B.S., Colorado State University, Mathematics

International Experience

  • Australia (population monitoring, invasive species, population estimation, damage estimation)
  • New Zealand (population indexing methods for brush-tailed possum)
  • Germany (population indexing)
  • Mexico (rodents, rabies baiting)
  • Morocco (ecotoxicology of insecticides)
  • Puerto Rico (invasive species, Puerto Rican parrot, rabies, monkey damage)
  • Guam (brown treesnakes)
  • Haiti (animal damage to crops)
  • Ethiopia (indexing wildlife)

Contact Information

Recent Publications


Baldwin, R.A., N. Quinn, D.H. Davis, and R.M. Engeman. 2014. Effectiveness of rodenticides for managing invasive roof rats and native deer mice in orchards.  Environmental Science and Pollution Research 21(9): 5795-5802.  doi: 10.1007/s11356-014-2525-4.

Engeman, R.M., T. Guerrant, G. Dunn, S.F. Beckerman, C. Anchor. 2014. Benefits to rare plants and highway safety from annual population reductions of a "native invader," white-tailed deer, in a Chicago-area woodland. Environmental Science and Pollution Research 21(2): 1592-1597. doi: 10.1007/s11356-013-2056-4.

Piaggio, A.J., R.M. Engeman, M.W. Hopken, J.S. Humphrey, K.L. Keacher, W.E. Bruce, and M.L. Avery. 2014. Detecting an elusive invasive species: a diagnostic PCR to detect Burmese python in Florida waters and an assessment of persistence of environmental DNA. Molecular Ecology Resources, 14: 374–380. doi: 10.1111/1755-0998.12180.

Allen, B.L., R.M. Engeman, and L.K-P. Leung. 2014. The short-term effects of a routine poisoning campaign on the movements and detectability of a social top-predator. Environmental Science and Pollution Research 21: 2178-2190. doi 10.1007/s11356-013-2118-7.

Engeman, R., T. Hershberger, S. Orzell, R. Felix, G. Killian, J. Woolard, J. Cornman, D. Romano, C. Huddleston, P. Zimmerman, C. Barre, E. Tillman, M. Avery. 2014. Impacts from control operations on a recreationally hunted feral swine population at a large military installation in Florida. Environmental Science and Pollution Research. Online. doi 10.1007/s11356-014-2727-9.

Berentsen, A.R., W.C. Pitt, J.D. Eisemann, R.M. Engeman. 2014. Longevity of rodenticide bait pellets in a tropical environment following a rat eradication program. Environmental Science and Pollution Research 21(3): 2283-2288. doi 10.1007/s11356-013-2148-1.


Hartmann, J.W., T.G. Guerrant, S.F. Beckerman, R.M. Engeman, and T.W. Seamans. 2013. Report to the Chicago Park District on conflicts with Ring-billed gulls and the 2013 Integrated Ring-billed Gull Damage Management Project. USDA, Wildlife Services. 29 p.

Ramey, C. A., K. W. Mills, J. W. Fischer, R. G. McLean, K.A. Fagerstone, and R. M. Engeman. 2013. Graphically characterizing the movement of a rabid striped skunk epizootic across the landscape in northwestern Wyoming. EcoHealth 10(3):246-256. doi: 10.1007/s10393-013-0853-3.

Allen B.L., Fleming P.J.S., Allen L.R., Engeman R.M.E., Ballard G., Leung L.K.P. 2013. As clear as mud: A critical review of evidence for the ecological roles of Australian dingoes. Biological Conservation 159:158-174. DOI: http://dx.doi.org/10.1016/j.biocon.2012.12.004.

Allen, B.L., L.R. Allen, R.M. Engeman, and L.K-P. Leung. 2013. Intraguild relationships between sympatric predators exposed to lethal control: predator manipulation experiments. Frontiers in Zoology 10:39. http://www.frontiersinzoology.com/content/10/1/39

Baldwin R.A., Marcum D.B., Orloff S.B., Vasquez S.J., Wilen C.A.W., Engeman R.M. 2013. The influence of trap type and cover status on capture rates of pocket gophers in California. Crop Protection 46: 7-12.

Engeman, R.M., K.J. Couturier, R.K. Felix Jr., and M.L. Avery. 2013. Feral swine disturbance at important archaeological sites. Environmental Science and Pollution Research. 20:4093-4098. DOI: 10.1007/s11356-012-1367-1

Engeman, R.M., W.C. Pitt, A.R. Berentsen and J.D. Eisemann. 2013. Assessing spatial variation and overall density of aerially broadcast toxic bait during a rat eradication on Palmyra Atoll. Environmental Science and Pollution Research Institute. 20:480–487. DOI 10.1007/s11356-012-1050-6.

Engeman, R.M., G. Massei, M. Sage, and M.N. Gentle. 2013. Monitoring wild pig populations : a review of methods. Environmental Science and Pollution Research 20(11): 8077-8091. doi 10.1007/s11356-013-2002-5.

Hartmann, J.W., S.F. Beckerman, T.W. Seamans, R.M. Engeman, and S. Beazley. 2013. Report to the City of Chicago on conflicts with ring-billed gulls and the 2012 integrated ring-billed gull damage management project. Chicago, USDA, Wildlife Services

Thomas, J.F., R.M. Engeman, E.A. Tillman, J.W. Fischer, S.L. Orzell, D.H. Glueck, R.K. Felix, and M.L. Avery. 2013. Optimizing line intercept sampling and estimation for feral swine damage levels in ecologically sensitive wetland plant communities. Environmental Science and Pollution Research 20:1503-1510. DOI 10.1007/s11356-012-1004-z.

Mathies, T., B. Levine, R. Engeman, and J.A. Savidge. 2013. Pheromonal control of the invasive brown treesnake: potency of female sexual attractiveness pheromone varies with ovarian state. International Journal of Pest Management, 59:2, 141-149. doi: 10.1080/09670874.2013.784374


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