Dr. Page Klug is the Supervisory Research Wildlife Biologist at the North Dakota Field Station.
Ph.D., Kansas State University, Biology, Interactions between grassland birds and their snake predators: the potential for conservation conflicts in the tallgrass prairie
M.S., University of Nebraska at Omaha, Biology, The effects of grassland patch and landscape characteristics on the activity and abundance of predators of grassland birds
B.S., Drake University, Environmental Science and Policy/Biology
Guam (brown treesnake ecology and control)
Costa Rica (impact of small mammals on tree diversity and recruitment)
Current Academic Affiliation:
Graduate Degree Program in Biological Sciences, North Dakota State University
USDA-APHIS-WS NWRC, North Dakota Field Station
Biological Sciences, Dept. 2715
North Dakota State University
PO Box 6050
Fargo, ND 58108-5060
Mahoney, J.L., W. Reed, G. Linz, and P. Klug. 2016. Female blackbirds' responses to stress during breeding: Possible implications for future management. Proceedings of the 27th Vertebrate Pest Conference 27:221-225.
Wandrie, L.J., W. Reed, M. Clark, K. Karels, P. Klug, and G. Linz. 2016. Behavioral responses of red-winged blackbirds to simulated predators: Evidence of potential habituation to video stimuli. Proceedings of the 27th Vertebrate Pest Conference 27: 253-257.
Other selected publications:
McEachern, MA, AA Yackel Adams, PE Klug, LA Fitzgerald, RN Reed. 2015. Brumation of introduced Tupinambis merianae (Squamata: Teiidae) in southern Florida. Southeastern Naturalist 14(2):319-328.
Klug, PE, RN Reed, FJ Mazzotti, MA McEachern, JJ Vinci, KK Craven, AA Yackel Adams. 2015. The influence of disturbed habitat on the spatial ecology of the black and white Argentine tegu (Tupinambis merianae), a recent invader in the Everglades ecosystem (Florida, U.S.A.). Biological Invasions 17(6):1785-1797
Cox, WA, LL Wolfenbarger, LI Berkeley, SE Engberg, W Janousek, PE Klug, N Rider, JP McCarty. 2014. Grassland bird communities on conservation and marginal grasslands in an agricultural landscape. Agriculture, Ecosystems & Environment 193:53-59.
Klug, PE, J Fill, KA With. 2011. Spatial ecology of Eastern Yellowbelly Racer (Coluber constrictor flaviventris) and Great Plains Ratsnake (Pantherophis emoryi) in a contiguous tallgrass prairie landscape. Herpetologica 67(4): 428-439.
Klug, PE, KA With, SM Wisley. 2011. Population genetic structure and landscape connectivity of the Eastern Yellowbelly Racer (Coluber constrictor flaviventris) in the contiguous tallgrass prairie of northeastern Kansas, USA. Landscape Ecology 26:281–294.
Klug, PE, SL Jackrel, KA With. 2010. Linking snake habitat use to nest predation risk in grassland birds: the dangers of shrub cover. Oecologia 162:803-813.
Klug, PE, LL Wolfenbarger, JP McCarty. 2010. Snakes are important nest predators of Dickcissels in an agricultural landscape. Wilson Journal of Ornithology 122(4):799-803.
Klug, PE, LL Wolfenbarger, JP McCarty. 2009. The nest predator community of grassland birds responds to agroecosystem habitat at multiple scales. Ecography 32:973-982.
Klug, PE, KA With, SM Wisely. 2009. Isolation and characterization of microsatellite loci in the Eastern Yellowbelly Racer (Coluber constrictor flaviventris) for use in landscape genetic studies. (Molecular Ecology Resources database accessions 37758–37769). http://tomato.biol.trinity.edu/. Cited as: Molecular Ecology Resources Primer Development Consortium et al. 2009. Permanent Genetic Resources added to Molecular Ecology Resources Database 1 May 2009–31 July 2009. Molecular Ecology Resources 9:1460-1559.